The Gap Analysis Project (GAP) Analytical Database represents a synthesis of three core datasets for the conterminous U.S. Specifically 1) the GAP/LANDFIRE National Terrestrial Ecosystems_2011; 2) the Protected Areas Database of the United States (PAD-US) 1.4; and 3) the Species Ranges and Habitat Distribution Models for all terrestrial vertebrates. This database provides a mechanism to efficiently obtain summary statistics of those for a variety of spatial extents, including US states, US counties, Landscape Conservation Cooperation Network Areas, EPA's Level III-IV Ecoregions of the United States, and Level I-III Ecoregions of North America and 12-digit (6th level) hydrologic units.